diff --git a/crates/zakura-chain/src/parameters/network.rs b/crates/zakura-chain/src/parameters/network.rs
index 38bfb1ca4..12a251d94 100644
--- a/crates/zakura-chain/src/parameters/network.rs
+++ b/crates/zakura-chain/src/parameters/network.rs
@@ -342,6 +342,27 @@ impl Network {
             .collect()
     }
 
+    /// Returns the first height at which version 4 transactions are invalid.
+    ///
+    /// [ZIP 2003] deprecates version 4 transactions at NU7. The
+    /// `nu7-experimental` feature must be enabled. A network that does not
+    /// activate NU7 keeps accepting them.
+    ///
+    /// [ZIP 2003]: https://zips.z.cash/zip-2003
+    pub fn v4_deprecation_height(&self) -> Option<Height> {
+        if cfg!(feature = "nu7-experimental") {
+            NetworkUpgrade::Nu7.activation_height(self)
+        } else {
+            None
+        }
+    }
+
+    /// Returns whether version 4 transactions are invalid at `height` on this network.
+    pub fn is_v4_deprecated(&self, height: Height) -> bool {
+        self.v4_deprecation_height()
+            .is_some_and(|deprecation_height| height >= deprecation_height)
+    }
+
     /// Returns the height at which the soft fork that temporarily disables Orchard
     /// actions in transactions activates, if it is configured for this network.
     pub fn temporary_orchard_disabling_soft_fork_height(&self) -> Option<Height> {
diff --git a/crates/zakura-consensus/src/transaction.rs b/crates/zakura-consensus/src/transaction.rs
index 460599930..25aa1aaf8 100644
--- a/crates/zakura-consensus/src/transaction.rs
+++ b/crates/zakura-consensus/src/transaction.rs
@@ -993,7 +993,7 @@ where
         let tx = request.transaction();
         let nu = request.upgrade(network);
 
-        Self::verify_v4_transaction_network_upgrade(&tx, nu)?;
+        Self::verify_v4_transaction_network_upgrade(&tx, network, request.height(), nu)?;
 
         let sapling_bundle = cached_ffi_transaction.sighasher().sapling_bundle();
 
@@ -1010,11 +1010,28 @@ where
         .and(Self::verify_sapling_bundle(sapling_bundle, &sighash, tx_id)))
     }
 
-    /// Verifies if a V4 `transaction` is supported by `network_upgrade`.
+    /// Verifies if a V4 `transaction` is supported by `network_upgrade` at `height` on
+    /// `network`.
     fn verify_v4_transaction_network_upgrade(
         transaction: &Transaction,
+        network: &Network,
+        height: block::Height,
         network_upgrade: NetworkUpgrade,
     ) -> Result<(), TransactionError> {
+        // # Consensus
+        //
+        // > [NU7 onward] The transaction version number MUST be 5 or 6.
+        //
+        // https://zips.z.cash/zip-2003
+        //
+        // ZIP 2003 deprecates V4 transactions at NU7.
+        if network.is_v4_deprecated(height) {
+            return Err(TransactionError::UnsupportedByNetworkUpgrade(
+                transaction.version(),
+                network_upgrade,
+            ));
+        }
+
         match network_upgrade {
             // Supports V4 transactions
             //
@@ -1039,7 +1056,8 @@ where
             | NetworkUpgrade::Nu6
             | NetworkUpgrade::Nu6_1
             | NetworkUpgrade::Nu6_2
-            | NetworkUpgrade::Nu6_3 => Ok(()),
+            | NetworkUpgrade::Nu6_3
+            | NetworkUpgrade::Nu7 => Ok(()),
 
             #[cfg(zcash_unstable = "zfuture")]
             NetworkUpgrade::ZFuture => Ok(()),
@@ -1047,8 +1065,7 @@ where
             // Does not support V4 transactions
             NetworkUpgrade::Genesis
             | NetworkUpgrade::BeforeOverwinter
-            | NetworkUpgrade::Overwinter
-            | NetworkUpgrade::Nu7 => Err(TransactionError::UnsupportedByNetworkUpgrade(
+            | NetworkUpgrade::Overwinter => Err(TransactionError::UnsupportedByNetworkUpgrade(
                 transaction.version(),
                 network_upgrade,
             )),
diff --git a/crates/zakura-consensus/src/transaction/tests.rs b/crates/zakura-consensus/src/transaction/tests.rs
index d73e21e36..7158680dc 100644
--- a/crates/zakura-consensus/src/transaction/tests.rs
+++ b/crates/zakura-consensus/src/transaction/tests.rs
@@ -46,7 +46,7 @@ use zakura_chain::{ironwood, orchard};
 
 use zakura_node_services::mempool;
 use zakura_state::ValidateContextError;
-use zakura_test::mock_service::MockService;
+use zakura_test::mock_service::{MockService, PanicAssertion};
 
 use crate::{error::TransactionError, primitives, transaction::POLL_MEMPOOL_DELAY, BoxError};
 
@@ -4452,6 +4452,93 @@ async fn v5_with_duplicate_orchard_action() {
     }
 }
 
+/// Checks that ZIP 2003 accepts V4 transactions below NU7 and rejects them from NU7.
+#[test]
+fn v4_deprecation_boundary() {
+    let _init_guard = zakura_test::init();
+
+    let nu7 = Height(2_000_000);
+    let tx = test_transactions(&Network::Mainnet)
+        .map(|(_, tx)| tx)
+        .find(|tx| matches!(**tx, Transaction::V4 { .. }))
+        .expect("V4 tx");
+
+    let activation_heights = ConfiguredActivationHeights {
+        before_overwinter: Some(1),
+        overwinter: Some(2),
+        sapling: Some(3),
+        blossom: Some(4),
+        heartwood: Some(5),
+        canopy: Some(6),
+        nu5: Some(7),
+        nu6: Some(8),
+        nu6_1: Some(9),
+        nu6_2: Some(10),
+        nu6_3: Some(11),
+        nu7: Some(nu7.0),
+    };
+
+    let at_nu7 = Parameters::build()
+        .with_activation_heights(activation_heights)
+        .expect("activation heights are valid")
+        .clear_funding_streams()
+        .to_network()
+        .expect("failed to build configured network");
+
+    assert_eq!(
+        at_nu7.v4_deprecation_height(),
+        cfg!(feature = "nu7-experimental").then_some(nu7)
+    );
+    assert!(
+        verify_v4_at(&at_nu7, &tx, nu7.previous().expect("height")).is_ok(),
+        "a V4 transaction must be valid below the deprecation height",
+    );
+    if cfg!(feature = "nu7-experimental") {
+        assert_eq!(
+            verify_v4_at(&at_nu7, &tx, nu7),
+            Err(TransactionError::UnsupportedByNetworkUpgrade(
+                4,
+                NetworkUpgrade::Nu7
+            )),
+            "a V4 transaction must be invalid at the deprecation height",
+        );
+    } else {
+        assert!(
+            verify_v4_at(&at_nu7, &tx, nu7).is_ok(),
+            "a default build must keep accepting V4 transactions",
+        );
+    }
+
+    // A network without NU7 keeps accepting V4 transactions.
+    let no_nu7 = Parameters::build()
+        .to_network()
+        .expect("failed to build configured network");
+
+    assert_eq!(no_nu7.v4_deprecation_height(), None);
+    assert!(!no_nu7.is_v4_deprecated(Height::MAX));
+}
+
+/// A [`Verifier`] with concrete service types, so a test can name its associated
+/// functions without the compiler inferring the services from a call.
+type TestVerifier = Verifier<
+    MockService<zakura_state::Request, zakura_state::Response, PanicAssertion>,
+    MockService<mempool::Request, mempool::Response, PanicAssertion>,
+>;
+
+/// Runs the V4 network upgrade check for `tx` at `height` on `network`.
+fn verify_v4_at(
+    network: &Network,
+    tx: &Transaction,
+    height: Height,
+) -> Result<(), TransactionError> {
+    TestVerifier::verify_v4_transaction_network_upgrade(
+        tx,
+        network,
+        height,
+        NetworkUpgrade::current(network, height),
+    )
+}
+
 /// Checks the activation boundary of the temporary Orchard-disabling soft fork:
 /// it is inactive below the configured height and active at and above it, can be
 /// disabled entirely, and Mainnet uses its fixed activation height.
diff --git a/docs/changelog/unreleased/856.md b/docs/changelog/unreleased/856.md
new file mode 100644
index 000000000..60c5440a1
--- /dev/null
+++ b/docs/changelog/unreleased/856.md
@@ -0,0 +1,7 @@
+## Changed
+
+- ZIP 2003 "Disallow version 4 transactions": The off-by-default
+  `nu7-experimental` feature makes Zakura reject version 4 transactions from
+  NU7 activation. This deprecates new Sprout transactions without exposing a
+  separate consensus configuration
+  ([#856](https://github.com/zakura-core/zakura/pull/856)).